It's just an aspect ratio, both should fit fine. The 120/90-18 has a slightly larger diamater (1in) than the 120/80-18. The 4.00x18 is a (decimal) 102/100-18 and is a tad smaller than the 120/80-18 in tire diameter.
Use this calculator:
http://www.csgnetwork.com/tireinfo4calc.html